The Autobots are under siege from foes both known and unknown. Jack tags along with Arcee on a routine patrol and the pair soon discover a crashed spaceship; the ship's occupant could be one of the most dangerous enemies the Autobots have ever encountered. Another crashed ship is discovered by Optimus Prime, this time with Autobot markings; what’s inside however could prove more harmful than helpful to his cause. And while the Autobots face a series of lethal threats, Megatron and Starscream continue to battle for supremacy over the Decepticons.